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Identification of a novel cis-regulatory element essential for immune tolerance

Identification of a novel cis-regulatory element essential for immune tolerance
Identification of a novel cis-regulatory element essential for immune tolerance
Thymic central tolerance is essential to preventing autoimmunity. In medullary thymic epithelial cells (mTECs), the Autoimmune regulator (Aire) gene plays an essential role in this process by driving the expression of a diverse set of tissue-specific antigens (TSAs), which are presented and help tolerize self-reactive thymocytes. Interestingly, Aire has a highly tissue-restricted pattern of expression, with only mTECs and peripheral extrathymic Aire-expressing cells (eTACs) known to express detectable levels in adults. Despite this high level of tissue specificity, the cis-regulatory elements that control Aire expression have remained obscure. Here, we identify a highly conserved noncoding DNA element that is essential for Aire expression. This element shows enrichment of enhancer-associated histone marks in mTECs and also has characteristics of being an NF-κB-responsive element. Finally, we find that this element is essential for Aire expression in vivo and necessary to prevent spontaneous autoimmunity, reflecting the importance of this regulatory DNA element in promoting immune tolerance.
0022-1007
1993-2002
Laflam, Taylor N.
36eea53d-4f28-4f67-b50e-537f2870fda6
Seumois, Grégory
0be7d3d6-5526-458c-aa5c-cce52410a2ed
Miller, Corey N.
684ddfaa-f2f3-442e-9a3c-41fb93a7a9a3
Lwin, Wint
5bfd6e9c-3233-4a39-b530-66e7335d8dd9
Fasano, Kayla J.
0926de00-4616-4a77-9967-7e84b2d70b2f
Waterfield, Michael
9f2c1bb7-ec85-44f5-855a-318e7984a246
Proekt, Irina
d7e66497-fe49-4458-9c4e-3a417be6a523
Vijayanand, Pandurangan
79514f33-66cf-47cc-a8fa-46bbfc21b7d1
Anderson, Mark S.
be48ded5-417a-4219-a563-f1c7b73c3609
Laflam, Taylor N.
36eea53d-4f28-4f67-b50e-537f2870fda6
Seumois, Grégory
0be7d3d6-5526-458c-aa5c-cce52410a2ed
Miller, Corey N.
684ddfaa-f2f3-442e-9a3c-41fb93a7a9a3
Lwin, Wint
5bfd6e9c-3233-4a39-b530-66e7335d8dd9
Fasano, Kayla J.
0926de00-4616-4a77-9967-7e84b2d70b2f
Waterfield, Michael
9f2c1bb7-ec85-44f5-855a-318e7984a246
Proekt, Irina
d7e66497-fe49-4458-9c4e-3a417be6a523
Vijayanand, Pandurangan
79514f33-66cf-47cc-a8fa-46bbfc21b7d1
Anderson, Mark S.
be48ded5-417a-4219-a563-f1c7b73c3609

Laflam, Taylor N., Seumois, Grégory, Miller, Corey N., Lwin, Wint, Fasano, Kayla J., Waterfield, Michael, Proekt, Irina, Vijayanand, Pandurangan and Anderson, Mark S. (2015) Identification of a novel cis-regulatory element essential for immune tolerance. The Journal of Experimental Medicine, 212 (12), 1993-2002. (doi:10.1084/jem.20151069).

Record type: Article

Abstract

Thymic central tolerance is essential to preventing autoimmunity. In medullary thymic epithelial cells (mTECs), the Autoimmune regulator (Aire) gene plays an essential role in this process by driving the expression of a diverse set of tissue-specific antigens (TSAs), which are presented and help tolerize self-reactive thymocytes. Interestingly, Aire has a highly tissue-restricted pattern of expression, with only mTECs and peripheral extrathymic Aire-expressing cells (eTACs) known to express detectable levels in adults. Despite this high level of tissue specificity, the cis-regulatory elements that control Aire expression have remained obscure. Here, we identify a highly conserved noncoding DNA element that is essential for Aire expression. This element shows enrichment of enhancer-associated histone marks in mTECs and also has characteristics of being an NF-κB-responsive element. Finally, we find that this element is essential for Aire expression in vivo and necessary to prevent spontaneous autoimmunity, reflecting the importance of this regulatory DNA element in promoting immune tolerance.

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Accepted/In Press date: 1 October 2015
e-pub ahead of print date: 2 November 2015
Published date: 16 November 2015

Identifiers

Local EPrints ID: 436481
URI: http://eprints.soton.ac.uk/id/eprint/436481
ISSN: 0022-1007
PURE UUID: e0a7738f-571a-4836-9ac7-6d8bc8b39eac
ORCID for Pandurangan Vijayanand: ORCID iD orcid.org/0000-0001-7067-9723

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Date deposited: 11 Dec 2019 17:30
Last modified: 16 Mar 2024 05:37

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Contributors

Author: Taylor N. Laflam
Author: Grégory Seumois
Author: Corey N. Miller
Author: Wint Lwin
Author: Kayla J. Fasano
Author: Michael Waterfield
Author: Irina Proekt
Author: Pandurangan Vijayanand ORCID iD
Author: Mark S. Anderson

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